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By van Jaarsfeld E.J., de Villiers P.U.

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The first phase, prophase to anaphase I, results in exchange of DNA between the pairs of chromosomes followed by their separation with both chromatids present. The second phase (metaphase II to telophase II) is a mitosis resulting in separation of the chromatids and the formation of four haploid cells. Related topics The cell cycle The plant cell (B1) Nucleus and genome (B5) Meristems and primary tissues (C1) Pollen and ovules (D2) Features of growth and development (F1) Cell division in plants occurs in meristems (Topic C1) and involves two parts: mitosis in which the chromosomes are replicated and sorted into two nuclei, and cytokinesis in which the cell wall, cytoplasm and organelles divide.

1). The apical meristem produces leaf primordia (which will form leaves) and bud primordia (which will form shoots). These are produced in position and order which gives rise to the characteristic form of the shoot which is recognizable for each species (see ‘architecture’ below). The shoot is surrounded by an epidermis. This outer layer provides the protective barrier between the stem and its environment and is covered in a lipid-based protective substance, cutin (Topics I1 and M4). Within the epidermis, cells of the ground tissues, the cortex, may be photosynthetic and occupy the space surrounding the vascular bundles.

The petals are usually the main organs attracting insects to flowers and many are brightly colored, though they may be photosynthetic as well. There may be an indefinite number of petals and/or sepals, but many plants have a fixed number, often three or five. The fertile parts of the flower are inside the petals (Fig. 1). They consist of the pollen-bearing male parts, stamens, in one or more whorls and the female carpels in the center. Stamens consist of a stalk, the filament, and an anther enclosing the pollen grains.

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